The nucleolus (also called nucleole) is a non-membrane bound structure found within the nucleus in which ribosomal RNA is transcribed, and is composed of protein and nucleic acids. The nucleolus ultrastructure can be visualized through an electron microscope while the organization and dynamics can be studied through fluorescent protein tagging and fluorescent recovery after photobleaching (FRAP). Malfunction of nucleoli can be the cause for several human diseases.
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